Showing posts with label goumoun. Show all posts
Showing posts with label goumoun. Show all posts

Sunday, 8 March 2009

Centaurism II

A while ago I discussed the goumoun, a six-limbed animal. I thought that freeing a pair of limbs for another purpose definitely occurred more than once in evolution, but that this principle sadly had no name. So I took it upon myself to call it 'centaurism'. Do not get me wrong: I am not claiming eternal fame for this invention, nor do I think evolution biology will suffer at all if it does not spread beyond this page.

Just treat it as an intellectual game. The original centaurs in Greek mythology had the body of a horse, but where its neck and head should be you would find the torso, arms and head of a human. Ancient Greeks were notoriously vague about the evolutionary history of their mythological beings, so we know very little about protocentauroids. Nothing, in fact. Did they walk on six limbs? The Greeks were probably more inclined towards a form of creationism: 'Poof!', leading to 'here's one I made earlier'.

On Furaha, the ancestors of neocarnivores did walk on six limbs in the typical hexapod fashion. There is nothing wrong with walking on four limbs, as tetrapods on Earth show, and indeed on Furaha. There was therefore ample room for evolutionary experimentation with front limbs in hexapod Furahan animals. They accordingly evolved into clubs, spears, lances, hatches and even nets (in the Microraptoria). If you want to see a few examples, simply travel to the land page on the Furaha site and have a look. Here's one of them:


Copyright Gert van Dijk

But centaurism is not confined to the goumoun or to Furaha. In fact, there are quite a few examples on Earth. Let's start with animals that started with four legs: are there any that stopped walking on all fours and got up on their hind limbs? Definitely; there are ostriches and other ratites, kangaroos, predatory dinosaurs and people, to name the most obvious.

What do they all do with their freed front legs? Some animals use them for grasping purposes at some times but to walk with at other times. Examples are the giant panda, lots of primates, as well as the kangaroo. If kangaroos move very slowly, they move on five limbs: the hind legs swing forwards while the body rests on the front legs and the tail.
Other animals do not appear to do very interesting things with their front legs, such as ostriches. I have seen a male ostrich flapping it to attract females, but that is about it. In fact, some Furahan analogues do exactly that, but, having freed not just two but four limbs, they look even odder (the Grec on the land page is a perfect example). The same probably goes for the front limbs of Tyrannosaurus. I know that its arms were less weak then they look, and am aware of theories saying the beast used its arms to help stand up from a lying position. But all in all these puny arms do not impress me at all. Faced with animals that do not do anything interesting with their freed front legs, perhaps the definition of centaurism should include a new non-locomotory purpose for the freed legs.

That thought reduces the list: primates are still in, as well as those predatory dinosaurs that used their front limbs to do something interesting with. That group goes under the name of Maniraptora. One of the most intriguing examples would be Therizinosaurus, here shown as seen in the BBC documentary 'The Giant Claw', a 'Walking with Dinosaurs special'. Not that much is known about the animal apart from its front legs with their huge claws, so the claws are to the right scale, but the rest is educated guesswork.

BBC Walking with Dinosaurs Specials: The Giant Claw
Click to enlarge

There is no reason to limit the principle of centaurism to animals starting with four walking limbs. Crabs are decapods ('ten-leggers'), but walk on eight legs. The front pair, the claws, are modified legs. In fact, the modifications are not that extensive, as is shown on an excellent animation on the internet from the University of Alberta and explaining how crabs' claws evolved and work. Here is a non-moving fragment to entice you to go and visit the original.

Click to enlarge

There are many other arthropod examples. Here is another one: Amblypygi or whip scorpions are spiderlike, but no longer walk on eight legs. Instead, one pair has evolved into long whiplike sensory organs. The next two photographs are from 'Life in the Undergrowth', another BBC documentary. It has some excellent footage of these animals. The whips are the thin limbs between the pedipalps and the walking legs. If you need more, check up on Uropygids in Wikipedia.

BBC Life in the Undergrowth
Click to enlarge

In conclusion centaurism is fairly common. The modified legs are front pairs in all cases. I have not found any examples of a hind pair of legs becoming centaurised. That is probably because of 'cephalisation': the front end of a body commonly has the most sensory organs, food goes in there, etc. Once you have cephalisation, any manipulatory limb is best placed at the front. But if anyone can think of a sound evolutionary way to free hind limbs from locomotion in an animal that is already thoroughly cephalised, AND that might fit on Furaha, I will consider making a sketch of one.

Sunday, 8 February 2009

The goumoun, or 'Centaurism I'

A while ago I discussed Glapumt'ians, a creature from a French series of bandes dessinées (comics) called 'Valérian et Laureline' by Christin and Mezières. It is time to return to their universe with the goumoun. Here he is, in the first frame he appears in. Note that he walks in a trot. Unlike Ralph the Glapumt'ian the goumoun only occurs in one album: 'Bienvenue sur Alflolol'. The story if that Galaxity, i.e., Earth's power centre, has turned the planet Technorog into a technological powerhouse, with extensive mines and factories. And then the original inhabitants come back in family groups. They turn out to live for some 16,000 years and left the planet for an outing some 4000 years ago. Now they wish to pick up life where they left it, on their home planet of Alflolol. The dumbfounded Earth people do not really know what to do. A such, the tale is fairly typical of the time it appeared: 1972, with echos of the 'military-industrial complex', the establishment', etc. Laureline immediately bonds with them, while Valérian hesitates between following her and more or less doing his job. The Alflololians like a good party, and manage to wreck the neatly managed planet by camping in the wrong places, going out hunting leaving a wake of wrecked installations, etc. The following illustration shows the cover of the 1974 Dutch version (In Dutch Valérian is called Ravian, probably because of associations with the tranquilizing herb 'valeriaan'). The 'fish' they are hunting (a furutz) doesn't like hydrodynamically sound, with all those protuberances, but this series isn't about biomechaniscal rectitude. He looks exciting!
Click to enlarge
The goumoun is the pet of one family of Alflololians; he is definitely relatively intelligent and from day one the goumoun and Laureline become the best of friends. As I noted before, this series makes no attempt to make alien species truly alien. The facial expressions on the goumoun's face are as easy for us to read as those of other people, or, come to that, of Glapum'tians! In the course of the story Laureline is carried away by a monster looking like a giant octopus brachiating through the jungle. Valérian is afraid of hitting Laureline by mistake with his impressive-looking space gun, but luckily the goumoun comes to the rescue. The thing I like about the goumoun is that he has four walking legs and two other legs that function a bit like arms. There doesn't seem to be a thumb, and we never see him actually pick up objects with them, which is a bit odd; why have arms if you do not use them as such. Then again, kangaroos don't do that much with their front legs, do they? Of course, many predators on Furaha have a similar arrangement of limbs, with four locomotory limbs and two front legs adapted for hunting. In their case, evolution started with a six-legged design. In time, the first pair of limbs lost its locomotor role and became free for other purposes. There is a page on such animals on the Furaha site (go to the land page), but here is one of the animals on it anyway: While thinking about the concept of freeing a frontal pair of legs from locomotion, it struck me that I was unaware of a name for this evolutionary trick. This event has occurred on Earth quite a few times on Earth, but does not seem to have a name. If there is indeed no name for it (and please tell me if I am wrong!) one must be invented: let's call it 'centaurism'. The goumoun is an example of centaurism, and so are the neocarnivores with their raptorial specialisations. I think centaurism is a worthy subject for a future essay, so I will get back to it. Later. Meanwhile, if you want to read the album, you may not be in luck if you are restricted to English, as there does not seem to be a translated version. But in France the series is an established success, and you can even buy a statuette of the goumoun: